Inspired by her grandfather's stories, young Shinko embarks on wild flights of fancy about the history of the little town she lives in. But dreams can't fend off the realities of growing up.

Upon reaching the train station to death, a dejected soul is informed that he is lucky and will have another chance at life. He is placed in the body of a 14-year-old boy named Kobayashi Makoto, who has just committed suicide. Watched over by a neutral spirit named Purapura, the soul must figure out what his greatest sin and mistake in his former life was, before his time limit in Makoto's body runs out. He also has a number of other lesser duties he must complete, such as understanding what led Makoto to commit suicide in the first place and learning how to enjoy his second chance at life.

A small, lone Kappa miraculously survives over 200 years into modern day Japan, when he is found by young Koichi. Coo, who is secretly adopted by Koichi's family, searches with his new human friend for unpopulated places in hopes of finding any remaining of his kind.

“东京决战”一年后，布里塔尼亚少年鲁路修在11区（原日本国）过着平凡的学生生活。但是，鲁路修与弟弟罗洛的一次出行，遇到了黑色骑士团的余党。在与少女C.C再次结成契约之后，尘封的记忆摆在了鲁路修的面前。
身为帝国王子的鲁路修，为了建立人人平等的世界、让妹妹娜娜丽幸福的世界，而使用GEASS，令人绝对服从的力量，带领黑色骑士团向帝国宣战。带上假面化名ZERO的他，却在一年前的“东京决战”中被好友朱雀击败，被帝国皇帝抹去了记忆。
现在，恢复记忆的鲁路修不仅要面对帝国的强大军事力量，更要面对虚假的弟弟罗洛、失踪的妹妹娜娜丽、不知敌友的中华联盟、内部出现分歧的黑色骑士团。面对内忧外患，鲁路修走上了GEASS之力的诅咒——孤独的王之路。