Writer and teacher Iris finds her comfortable, solitary New York life thrown into disarray after her closest friend and mentor bequeaths her his beloved 150 lb. Great Dane. The regal yet intractable beast, named Apollo, immediately creates practical problems for Iris, from furniture destruction to eviction notices, as well as more existential ones. Yet as Iris finds herself unexpectedly bonding with Apollo, she begins to come to terms with her past, and her own creative inner life in this story of healing, love, and friendship.

在天才哥哥艾伦（麦克思·明格拉 Max Minghella 饰）的阴影下，艾吉拉（弗劳拉·克罗斯 Flora Cross 饰）觉得自己简直没有一点长处，父亲索尔（理查·基尔 Richard Gere 饰）和母亲米利亚姆（朱丽叶·比诺什 Juliette Binoche 饰）显然更喜欢的是聪明伶俐的艾伦，而在他们之间，亦有着根深蒂固无法解决的问题。
在一次拼字大赛上，艾吉拉意外的展现出了令人惊叹的天赋，在赢得比赛的同时，她也获得了走向世界的机会。可是，对于艾吉拉的成功，一家人并没有表现出应有的高兴与自豪，因为随着艾吉拉的出名，一个埋藏在家族中多年的秘密即将被公之于众。眼看着家庭陷入了危机之中，艾吉拉明白，她必须依靠自己的力量，重新将家人凝聚在一起。