"Jellicle" cats join for a Jellicle ball where they rejoice with their leader, Old Deuteronomy. One cat will be chosen to go to the "Heavyside Layer" and be reborn. The cats introduce themselves.

Mia, an aspiring actress, serves lattes to movie stars in between auditions and Sebastian, a jazz musician, scrapes by playing cocktail party gigs in dingy bars, but as success mounts they are faced with decisions that begin to fray the fragile fabric of their love affair, and the dreams they worked so hard to maintain in each other threaten to rip them apart.