Searching for a safer habitat, the prehistoric Crood family discovers an idyllic, walled-in paradise that meets all of its needs. Unfortunately, they must also learn to live with the Bettermans -- a family that's a couple of steps above the Croods on the evolutionary ladder. As tensions between the new neighbors start to rise, a new threat soon propels both clans on an epic adventure that forces them to embrace their differences, draw strength from one another, and survive together.

A young teenager named Mikey Walsh finds an old treasure map in his father's attic. Hoping to save their homes from demolition, Mikey and his friends Data Wang, Chunk Cohen, and Mouth Devereaux run off on a big quest to find the secret stash of Pirate One-Eyed Willie.

故事发生在上世纪的三十年代，路易斯（凯莉·霍威 Keeley Hawes 饰）的丈夫意外去世，留下了四个孩子格兰特（米罗·帕克 Milo Parker 饰）、劳伦斯（乔什·奥康纳 Josh O'Connor 饰）、莱斯利（卡伦·伍德豪斯 Callum Woodhouse 饰）、玛格（黛西·沃特斯通 Daisy Waterstone 饰），以及一笔遗产。如今，这笔遗产已经被花的差不多了，孩子们亦在渐渐长大，问题日益凸显，使路易斯感到力不从心。 在巨大的压力之下，路易斯决定带领一家人远赴生活成本较小的希腊科孚岛定居，希望在那里开始新的生活。然而她的决定显然没有受到孩子们的欢迎和赞成，更何况何科孚岛甚至都没有通电。路易斯这个本想修复家庭关系的决定，却反而让兄弟姐妹们之间火药味十足。