坐落于某个小镇的兔山商店街（うさぎ山商店街）上，有个经营饼店的少女－－北白川玉子。除了帮忙店里的事情，她亦有跟学校的朋友常盘绿和牧野神奈参与舞棒部活动。而饼店对面的大路家是同一行业的竞争对手，两边的父亲经常都因生意问题而喧闹一番。然而大路家的儿子大路饼藏却是玉子的朋友，但玉子从没注意到饼藏的好感。总而言之，玉子在商店街众人的护荫下，热闹地度过每一天，生活可谓过得相当快乐。

除夕，也就是玉子的生日，处于商店街年末最繁忙的时刻。庆祝仪式已是商店街上下的惯例，唯独饼藏今年没有准时来到。直到他带着礼物来到之时……

商店街突然出现了一只懂得说人话的鸟，犹如飞车般降临到众人面前。偶然变得越来越热闹的兔山商店街，亦正式开始其稍微不同的新一年。

2007年的热门动画《幸运星》以OVA形式推出了续篇。御宅族女孩泉此方、性格迥异的柊家姐妹、经常考虑问题不周到的高良美幸，四位女孩又要开始新的故事了。
小狗Cherry的奇怪举动搞得大家哭笑不得；与老师一起玩网络游戏引出了话题大讨论；姐姐柊镜突然进入不伦不类的灰姑娘的世界；排球比赛中妹妹柊司努力挑战自我，要做的比姐姐还好；去山区郊游的四人陷入了迷路的困境；放学后大家一起去的宠物店似乎隐藏着不可告人的秘密。
融合了日常生活与动漫恶搞的《幸运星》故事又开始了。

Kumiko and her teammates have won the Kyoto regional tournament. From now on, life will be getting a lot more hectic. With the national tournament fast approaching, Kumiko's band is doubling down on practice and discipline, leaving little time for sleep or other recreational activities. The redoubled practice schedule brings Kumiko and Reina closer together, but elsewhere in the band, it seems like drama threatens to tear the group's unity apart. The ghost of the prior year's mass band exodus still haunts many of Kumiko's classmates, and with former band member Nozomi Masumi now asking to return, it seems like the grudges of the past will soon be impossible to ignore.

Spring in the first year of high school. Kumiko, a member of the brass band in junior high school, visits the high school brass band club with classmates Hazuki and Sapphire. There, she comes across Reina, her former classmate from junior high. Hazuki and Sapphire decide to join the club, but Kumiko can’t make up her mind. She recollects her experience with Reina at a competition in junior high school.