When her father unexpectedly passes away, young Ella finds herself at the mercy of her cruel stepmother and her daughters. Never one to give up hope, Ella's fortunes begin to change after meeting a dashing stranger in the woods.

亚茨拉斐尔（迈克尔·辛 Michael Sheen 饰）是守卫城门的天使，而克劳利（大卫·田纳特 David Tennant 说）则是堕落的恶魔。两人几千年来都一直隐姓埋名生活在人间，并且通过不断的尝试，在天堂和地狱之间建立了井水不犯河水的互惠互利的关系。
克劳利奉撒旦之命，将婴儿敌基督送到修道院，同美国外交官的新生儿做交换，当敌基督长到11岁时，就是世界末日来临之时。为了防止世界被毁灭，亚茨拉斐尔和克劳利决定通过影响这个孩子的善恶观念来避免世界的终结。在努力了11年之后，两人震惊的发现，他们竟然弄错了对象，而真正的敌基督此时生活在一个名为塔德菲尔德偏僻小镇中。

As the war between the vampires and the Lycans rages on, Selene, a former member of the Death Dealers (an elite vampire special forces unit that hunts werewolves), and Michael, the werewolf hybrid, work together in an effort to unlock the secrets of their respective bloodlines.